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Topic. Compound Eyes Of. Contents. Receptors in Insects Photoreception in Insects Compound Eyes Ommatidia Components of Ommatidia Photochemistry of Insect vision Forms of Compound Eyes Significance of Compound Eyes Color vision in Some Orders Of Insects Summary
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Topic Compound Eyes Of Entomology
Contents • Receptors in Insects • Photoreception in Insects • Compound Eyes • Ommatidia • Components of Ommatidia • Photochemistry of Insect vision • Forms of Compound Eyes • Significance of Compound Eyes • Color vision in SomeOrders Of Insects • Summary • References
Receptors in Insects • Mechanoreceptors • Auditory receptors • Stretch receptors • Chemoreceptors • Olfactory receptors • Gustatory receptors • Thermo receptors • Photo receptors
Vision is the perception of light.Roles of vision in insects • Photoreception in Insects Entomology
Photoreception in Insects • Photoreceptors:1. Ocelli. 2. Stemmata3. Compound eyes
Dorsal ocelli • In larvae of hemimetabolous insects and in nearly all adults. • Poor perception of form. • Active in orientation to a light source. Ref.02
Stemmata • In larvae of holometabolous insects • Do not produce clear images • Most caterpillars can discriminate some shapes and they can orientate themselves with respect to boundaries.
Compound Eyes: • Most adult insects have a single pair of compound eyes. • Reduced or absent in parasitic forms, many soil insects, and in some species that live in very dark places.
Facet: Hexagonal Components of Compound Eye. Entomology
Ommatidia • Basic unit of compound eyes • Vary in size and number. • Honey bee has 4900 Ommatidia in Queen, 6300 in workers and 13000 in donors. • Pomera punctatissima have only one ommatidium in each eye. • The sizes of Ommatidia vary from about 5 to 40 microns in diameter . Ref.02
Components of Ommatidia • Optical parts: 1. Corneal lens 2. Crystalline cone • Sensory parts: 1. Retinula cells 2. Rhabdom • A nerve axon projects from each retinula cell. • Optic nerve (YADAV. M) Ref.01
Photochemistry of Insect vision • Photons are caught on the rhabdome by retinal. • Retinal is connected to opsin, forming rhodopsin. • On absorption of photon, retinal changes its form from bent to straight. • When retinal changes its form, it separates from the rhodopsin and the opsin triggers a nerve cell. • The nerve cells conduct the signal to the brain. Ref.02
Forms of Compound Eyes • Apposition eyes • 2. Superposition eyes. Ref.01
Significance of Compound Eyes • Flicker effect The compound eye is excellent at detecting motion. As an object moves, ommatidia are turned on and off. Ref.03
Significance of Compound Eyes 2. Distance Perception • Most insects must be able to judge distance. • As in prey catching insects, in Grass hopper’s jumping, and when they are landing. • Simultaneous stimulation of ommatidia. Ref.01
Significance of Compound Eyes 3. Color vision • Some insects are able to distinguish colors • Most flower visiting insects exhibit preferences for blue and yellow. • Important in feeding and in court ship behavior
3. Color vision Bichromatic Insects. One pigment absorbs green and yellow light (550 nm); the other absorbs blue and ultraviolet light (<480 nm). Trichromatic Insects. Absorption maxima at 360 nm (UV), 440 nm (blue-violet), and 588 nm (yellow)
Human Vision Vs. Insect Sight • Colors visible to insects is higher in frequency (lower in wave length) • Violet light is the highest frequency of color humans can detect, but many insects can see a higher frequency of light invisible to us, ultraviolet light. REF 04
Ultraviolet vision Entomology
1. Vision in Order Hymenoptera • Eastern honey bee (Apis cerana). • Family: Apidae • The bees are red-blind. • “orange, yellow, and green are the same color” – yellow. • Sensitive to UV light. • They see blue colors best. • Sense of polarization of visible light in the sky. Ref.05
2. Vision in Order Odonata • Dragonfly(C. aenea) or Downy emerald • Family: Corduliidae • Compound eyes with wide-angle vision • Their eyes do not move. • 30,000 eye ommatidia • Orange to ultraviolet (UV) light. • Able to estimate distance based on the • distance between their eyes. Ref.06
3. Vision in Order Diptera • House flies (Musca domestica) • Family: Muscidae • The vision of the housefly is blurred • Compound apposition eyes • Can sense rapid motion approaching 200 • cycles per second. Ref.06
Vision in Mosquito • Mosquito (Culiseta longiareolata) • Family: Culicidae • Mosquitoes are attracted to black and to dark colors. (Howlett 1910) • Prefer the corners of a three-dimensional target. (Brown & Bennett) Ref.07
4. Vision in Order Coleoptera • Beetle (Tetraopes tetrophthalmus) • Family: Cerambycidae • Have infrared detection systems for night vision, fire detection, and other functions as to sense forest fires. • Beetles have refraction superposition eyes Ref.06
5. Vision in Order Blattaria American cockroach (Periplaneta americana), Family: Blattidae 2000 individual lenses Has receptorsfor UV light and green light. Ref.08
6. Vision in Order Homoptera AphidsSuper family: AphidoideaContains green, blue, and ultraviolet photoreceptors Ref.09
7. Vision in Order Lepidoptera • Butterfly(Papilio rutulus) • Family:Papilionidae • Can See simultaneously in every direction. • Trichromatic vision • It can distinguish night from the day • Can perceive colors in a high frequency (from 310 nm to 700) Ref.10
Vision in Order Lepidoptera • Gipsy Moth (Lymantria dispar). • Family: Lymantriidae • Can see in the back of their heads. • Neural summation • Have three types: blue, green, and ultraviolet. • Ultraviolet light reflection. Ref.12
Summary • Compound Eyes of Insects
References • YADAV. M, "Physiology of Insects," Discovery Publishing House New Delhi-110002, pp. 322-345 • http://entomology.unl.edu/ent801/ent801home.html • http://users.rcn.com/jkimball.ma.ultranet/BiologyPages/C/CompoundEye.html • www.cals.ncsu.edu/course/ent425/tutorial/colorvision.html • http://www.sewanee.edu/Chem/Chem&Art/Detail_Pages/ColorProjects_2003/Crone • www.eyedesignbook.com/index.html • http://www.ent.iastate.edu/dept/research/vandyk/hostseek.htm
References 8.http://www.sciencemag.org/cgi/content/abstract/171/3977/1254 9.http://www.ncbi.nlm.nih.gov/pmc/articles/PMC1945073/figure/pbio-0050187-g003 10. http://www.butterflyzone.org/butterfly-articles/butterfly-uv-vision.shtml 11.http://indianapublicmedia.org/amomentofscience/insect-color-vision 12.http://www.animalcorner.co.uk/insects/moths/moth.html
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